BOONE, N.C. — Josh Vaughan ran for 133 yards and three touchdowns to help Richmond upset defending national champion Appalachian State, 33-13, yesterday in the Football Championship Subdivision playoffs.

The Spiders (11-3) advance to the FCS — formerly Division 1-AA — semifinals for the second consecutive year. Richmond will play at New Hampshire next weekend.

Armanti Edwards threw for 323 yards and one touchdown, and ran for another score for the three-time FCS champion Mountaineers (11-3). But Edwards also threw a career-worst five interceptions in the second half, leading to four Richmond scores.

Eric Ward also had a rushing score, and Andrew Howard and Brian Radford both added field goals for the Spiders.

Appalachian State took a 7-0 lead late in the first quarter, driving 95 yards — aided by a 15-yard face mask call, and a pass interference penalty in the end zone on Richmond — before Edwards scored on a 2-yard run with 2:58 left.

The Spiders had a chance to tie the game with Vaughan scoring on a 13-yard run with 4.2 seconds left in the first quarter to pull the score to 7-6. Howard bounced his extra-point attempt off the right upright.

Richmond then took the lead on Howard's 36-yard field goal with 9:05 left in the third quarter, with back-to-back interceptions by Seth Williams to set up TD runs of 1 yard by Vaughan and 5 yards by Ward to put the Spiders ahead 23-7.

The Mountaineers pulled within 23-13 on Edwards' 4-yard touchdown pass to T.J. Courman with 1:51 left in the third quarter, but Richmond took advantage of two more interceptions to add a 27-yard field goal by Radford and a 1-yard Vaughan TD run.

NORTHERN IOWA 36, NEW HAMPSHIRE 34

CEDAR FALLS, Iowa — Pat Grace ran for two touchdowns and Terrell McMoore returned an interception 36 yards for what proved to be the decisive TD, lifting the third-seeded Panthers (12-2) over the Wildcats (11-3).

The Panthers survived a 100-yard interception runback for a touchdown by New Hampshire's John Clements to move into next weekend's semifinals.

New Hampshire's R.J. Toman was 16 of 30 passing for 243 yards and a touchdown, but he was intercepted four times and the Wildcats lost two fumbles.

JAMES MADISON 31, VILLANOVA 27

Harrisonburg, Va. — Rodney Landers threw for three touchdowns and scored on a 1-yard run with 1:38 to play as the top-seeded Dukes (12-1) beat the Wildcats (10-3) to advance to the semifinals against fifth-seeded Montana.

The Dukes trailed only once, after Matthew Szczur's 13-yard touchdown pass to Stephen Pimm with 6:54 left. But Landers drove James Madison 54 yards for the go-ahead touchdown and Marcus Haywood intercepted Chris Whitney to end Villanova's comeback hopes.

The game was a rematch of a regular-season contest won, 23-19, by James Madison on Landers' 35-yard Hail Mary touchdown pass on the final play.

MONTANA 24, WEBER STATE13

MISSOULA, Mont. — Chase Reynolds ran for 185 yards and two touchdowns as the host Grizzlies (13-1) avenged their only loss of the season with a victory over the Wildcats (10-4) in the quarterfinals.

Reynolds' final score, a 12-yard run with 4:02 remaining, came after Weber State (10-4) pulled to 17-13 on a 22-yard field goal by Jon Williams with just over nine minutes left.

Weber State quarterback Cameron Higgins, a Saint Louis alum from Hawai'i, passed for 270 yards and was intercepted once. The Wildcats also lost two fumbles.